I would say almost certainly more supple sidewalls.

Marathons are designed for the long distance tourer who needs maximum
flat resistance and overall durability.  Schwalbe does a good job
making the tire pleasant, but the puncture resistance stuff
incorprated in the tire tends to be more rigid and has less rolling
resistance.

If anything a wider tire with supple rubber will be more comfortable
with minimal impact on speed and acceleration.

On Jun 1, 9:26 am, Seth Vidal <skvi...@gmail.com> wrote:
> I've been riding an atlantis with schwalbe marathons - in their 40mm
> size. They've been nice and cushy if I let enough air out but have
> felt 'sluggish' at times and when I pump them up to max pressure they
> were a bit harsh. I decided to pickup a pair of the pasela panaracer
> TG - folding bead in 32mm. My LBS had them quite inexpensive at
> $30/each. Popped them on, pumped them up to about 75-80psi and went
> for a couple of rides.
>
> Things I noticed:
>  1. the atlantis feels a fair bit zippier.
>  2. the ride doesn't feel significantly rougher with the smaller tires.
>  3. I need  to lower the fenders to have them fit better :)
>
> So now what I'm wondering is:
>  1. Is the ride improved b/c the width is smaller?
>  2. is the ride improved b/c of the more supple sidewall?
>  3. Anyone else with similar experiences?
>
> Thanks
> -sv
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