I don't know specifics, but I do know that Mail uses more header criteria to 
group messages by conversation than by thread. I would imagine it takes the 
domain-related headers into account, but this is just a semi-educated guess. 
You can, for example, change the subject line in the middle of a thread, and it 
is still grouped into the original conversation.

Teresa
On Aug 6, 2011, at 11:14 AM, Paul Erkens wrote:

> Hi Teresa, Interesting, the difference you mention between threads and 
> conversations. When mail organizes messages by either, what are the 
> differences in criteria that mail relies on to do the job? Do you happen to 
> know? I'm asking because it looked like you do. See message below.
> Paul.
> On Jul 27, 2011, at 3:31 PM, Teresa Cochran wrote:
> 
>> That's the neat thing about conversations versus threads: they rely on more 
>> than the subject line, so that in most cases, you can change it, and still 
>> have messages within the conversation.
>> 
>> Teresa
>> On Jul 27, 2011, at 6:24 AM, Les Kriegler wrote:
>> 
>>> With the improved functioning of threads in Lion, it occurs to me that 
>>> valid subject lines are going to be more important than ever now.  BTW, I 
>>> just changed the subject line to reflect what I wished to discuss.
>>> 
>>> Les
>>> 
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