I can't answer your main question (about hyphenation after an apostrophe), but there are some points I can explain.

Le 16/10/2010 11:47, Cyril Niklaus a écrit :
In making the small version I include here, I also noticed something 
surprising: the hyphenation changed depending on the length of the text I cut 
*after*. Leaving one sentence or two did not give the same results. Cutting 
immediately after the sentence gave the correct l'in-formation and  
l'infor-mation was what I got when cutting before De toute part.

That's absolutely normal, that's even the reason why we use TeX :)
TeX builds a paragraph as a whole; if you remove some words at the end of your paragraph, it might change its entire shape.

I also noticed that including or not
\defaultfontfeatures{Mapping=tex-text}
changes things quite a bit while \frenchspacing did nothing obvious. I thought 
it would deal with spaces around the guillemets etc. but no. I'm wondering why 
I bothered including it. Is that a benefit from polyglossia?


\frenchspacing has nothing to with polyglossia, and it /is/ extremely important, even though you might not notice at once. It's a macro inherited from plain TeX, whose effect is to disable extra space after strong punctuation marks (e.g. a period), which extra space is used in (some flavors of) English typography. So keep it, although indeed it doesn't deal with space around guillemets.

Best,
Paul

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