A setup that *requires* from a recipient/sender to have a MX record is *broken*. Period. Talk to Wietse Venema if you disagree :)
Be that as it may in opinion, it was my observation that having this limitation has resulted in extremely minimal pushback with extremely significant gain. Everything we do is a trade off. If I could have a rule that had a 100% rate of hitting spam and 0% rate of hitting ham, I'd certainly love that up front. But the number of senders that need to send but don't want to receive bounces is actually pretty low in practice. The number of recipients without MX records that intend to receive mail have been zero across our entire platform, and we're not dealing with a small amount of email. In the recipient case, 100% have been registration form spam, causing junked up mail queues as they're all waiting for a DNS propagation event that isn't going to happen.
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