I had the same problem.  Although GAE supports 'reply_to' its not 
implemented in web2py.

I inserted two lines in web2py/gluon/tools.py; here is what the code looks 
like now:

            elif self.settings.server == 'gae':
                xcc = dict()
                if cc:
                    xcc['cc'] = cc
                if bcc:
                    xcc['bcc'] = bcc
                if reply_to:
    xcc['reply_to'] = reply_to

The elif is at line 594 in the current version 1.99.7

Seems to work fine now.

On Sunday, February 13, 2011 8:56:16 PM UTC-5, Richard Penman wrote:
>
> I have also experienced problems with the reply to field. Some info here:
> http://code.google.com/p/googleappengine/issues/detail?id=2802
>
> Also there are the restrictions on who you can set as the sender which you 
> probably already know about:
> http://code.google.com/appengine/docs/python/mail/sendingmail.html
>

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