You can do this by building a list of fields and then using the python 
unpack operator.  For example:

field_list = [db.auth_user.first_name]
if ask_for_lastname:
    field_list.append(db.auth_user.last_name)
rows = db(query).select(*field_list)

See 
https://docs.python.org/2/tutorial/controlflow.html#unpacking-argument-lists

-- 
Resources:
- http://web2py.com
- http://web2py.com/book (Documentation)
- http://github.com/web2py/web2py (Source code)
- https://code.google.com/p/web2py/issues/list (Report Issues)
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