After another look at the code, this does still seem to be deliberate,
if still counterintuitive. When drawing the header selection popup,
there's a comment that reads:
// if we currently only show one of either sender/receiver column
// modify the popup text in the way, that it displays the text of the other
of the two
Another strange comment later on:
// if we change the visibility of the receiver column,
// the sender column has to show either the sender or the receiver
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Kmail message list column context menu inconsistencies
https://launchpad.net/bugs/56603
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