Yep, have exactly same  issue.

As I suspected from point 0, this is the gnome-settings daemon crash
(rather hang more correctly).

To 'fix' the problem temporarily I did:

# the SIGKILL is importaint, because the program seems to be hung completely 
and ignores SIGTERM
killall -9 gnome-settings-daemon
gnome-settings-daemon.

#At that point panel looks ok, but nautilus not, so
#Then restart nautilus as it seems not to pick the change
killall nautilus

I bet that this is a race condition just happened to be exposed by slow
virtual CPU.

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https://bugs.launchpad.net/bugs/728803

Title:
  Ubuntu theme switches from partially broken to gnome default on
  VirtualBox guest

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