I noticed the same behavior in Totem. I found a related Gnome bug that explains 
this behavior:
https://bugzilla.gnome.org/show_bug.cgi?id=370732

Space is reserved to activating widgets for accessibility purposes. It can't be 
used for anything else. There's an entry in the Totem FAQ about this:
http://projects.gnome.org/totem/#faq

You can make Space toggle playback right now by first highlighting the
Play button in the toolbar by pressing Tab.

The behavior you're seeing during playback - the song starting from the
beginning - is cause by the action associated with the song list widget,
which is normally the selected widget. The behavior of the song list
widget could be changed to toggle playback. Or keyboard focus could
switch to the play button automatically after you start a song, though I
don't think that's a good idea.

** Bug watch added: GNOME Bug Tracker #370732
   https://bugzilla.gnome.org/show_bug.cgi?id=370732

-- 
Pressing Space key should toggle play/pause (pause when playing; play when 
paused)
https://bugs.launchpad.net/bugs/185928
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