Your code did not run on my computer, however the "atop" function should do
what you are looking for I guess. This is more or less your axis, only
changed a bit in your formula, think it looks better this way:
e.g.
par(mar=c(5,7,.5,.5), las=1, adj=.5, cex.lab=1.5)
plot(1, type="n"
, xlab="Week"
, ylab=expression(atop("Mean Respisration Rate",
umol%.%L^-1%.%g~(AFDM)^-1) ))
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