Here's just a ha'penny more, regarding the perplexing ankles of birds.

In our ankle, the long bone of the lower leg (tibia) articulates with several 
small bones, the tarsals, in the rear part of our plantigrade foot. The basic 
anatomy is similar in other mammals, except that in more cursorial forms, like 
cats and dogs, the rear part of their digitigrade feet (composed of tarsals and 
metatarsals) is generally raised above the ground, so that the ankle appears as 
an elevated, backward-pointing hinge, as in birds.

In Theropod dinosaurs, such as birds, the ankle joint forms among the tarsal 
bones, with some tarsal components fusing with the tibia (forming the 
tibiotarsus, above the ankle), and others, on the far side of the joint, fusing 
with each other and with metatarsal bones (forming the tarsometatarsus, below 
the ankle).

Clearly, the apparent length of a bird's leg is influenced not only by the 
actual lengths of these two bones, but also by their angle relative to each 
other, and the degrees to which the top of the tibiotarsus is concealed within 
the body plumage and the bottom of the tarsometatarsus (i.e., foot) is 
concealed by grass, water, or mud. The surest bet is to patiently assess the 
length of the whole tarsometatarsus when both the ankle and the toes are 
visible.

That said, the overall setup tends to look rather different in Dunlin and 
Curlew Sandpiper:

https://picasaweb.google.com/109808209543611018404/CalidrisII#5747741452480409570

Shai Mitra
Bay Shore, NY
________________________________
From: [email protected] 
[[email protected]] on behalf of Paul R Sweet 
[[email protected]]
Sent: Saturday, July 18, 2015 6:31 PM
To: robert adamo; NY BIRDS
Subject: Re: [nysbirds-l] A hummer of a breakfast + my "2 cents" !

Hi Robert

Just a quick note on bird anatomy. The lower (distal) long bone of the hindlimb 
to which the toes attach is the tarsometatarsus which is homologous with our 
foot. Thus the joint above this is the ankle not the knee. (look at the 
direction of hinging). The second long bone is the tibiotarsus which in 
shorebirds is partially exposed and scaly. The knee is where this joins the 
femur and is not visible as it is surrounded by muscle.

Happy shorebirding. Paul




From: robert adamo <[email protected]<mailto:[email protected]>>
Reply-To: robert adamo <[email protected]<mailto:[email protected]>>
Date: Saturday, July 18, 2015 at 1:17 AM
To: "[email protected]<mailto:[email protected]>" 
<[email protected]<mailto:[email protected]>>
Subject: [nysbirds-l] A hummer of a breakfast + my "2 cents" !

Over breakfast, my doctor (Michael Dempsey) called with good news, and I in 
turn, was able to share my good news re: the Ruby-throated Hummingbird, who was 
having her breakfast around the patio, while we were talking !

And now for my "2 cents" concerning the ID question posed by the Arie 
Gilbert/Phil Urubaru posting on 7/16. After reading Angus Wilson's response to 
them, in particular re: the longer legs of the Curlew Sandpiper, I would like 
to advance the idea that in some cases of limited viewing, just the length of 
the tibia is a fairly good mark in helping to separate the Curlew from the 
Dunlin. Admittedly, I can't find this referenced in the text of any of the many 
general bird guides, or the 2 excellent shorebird guides I own (the size is 
probably too variable), but a number of them show it in their illustrations. 
Thus, I know it stands to reason, one shouldn't rely on this, but if, and when, 
the next opportunity arises to consider it..."check it out" !  Although I 
certainly could not tell by looking at the Gilbert/Urubaru photos which of the 
two species the bird was, I can attest to identifying both species in the past, 
using the tibia, along with some of both species other field marks, ie, on 
10/3/78 at the East Pond, J.B.W.R.(my first C.S. sighting). "Winter plumage - 
saw white rump both while standing & in flight - noticed slight curve 
throughout length of bill - also noticed longer legs than Dunlin, with more 
showing above knee"...didn't know it was called tibia back then ! I have also 
been able to make this direct comparison on at least one other occasion.

Cheers,
Bob
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