New paper published titled “Accounting for sampling bias reveals a decline in 
abundance of endangered false killer whales in the main Hawaiian Islands”

Full citation: Badger, J.J., R.W. Baird, D.S. Johnson, A.L. Bradford, S.D. 
Mahaffy, M.A. Kratofil, T. Cullins, J.J. Currie, S.H. Stack, and E.M. Oleson. 
2025. Accounting for sampling bias reveals a decline in abundance of endangered 
false killer whales in the main Hawaiian Islands. Endangered Species Research 
57:325-340https://doi.org/10.3354/esr01423.

This is an open-access paper available from 
https://www.int-res.com/abstracts/esr/v57/esr01423 or from our Hawai‘i 
publication page https://cascadiaresearch.org/hawaii_publications/


ABSTRACT: We estimated abundance of the endangered main Hawaiian Islands (MHI) 
insular population of false killer whales Pseudorca crassidens from 1999–2022 
using a modeling technique that incorporates animal availability in a 
capture–recapture analysis. The population was sampled using different sampling 
methods, resulting in yearly encounter histories of 265 individuals and 53 
satellite-tagged whales. Survey effort and animal location data were separately 
analyzed using ker­nel density estimators, and the degree of overlap between 
these 2 processes was used to model detection probability in a Bayesian 
Jolly-Seber population model. This approach better addresses spatiotemporally 
variable sampling effort than traditional capture–recapture methods, improving 
the estimation of reliable abundance trends. Using simulated data, the model 
was robust to many sampling and ecological complications, such as variable low 
detectability, unequal tag deployment lengths, and variable social group sizes. 
Fitting the model to the MHI false killer whale data set, we found that the 
insular population of false killer whales remains small, with an estimated 139 
individ­uals (95% credible interval, CRI = [114, 162]) in 2022. The population 
appears to be in decline throughout the study period, with a mean annual 
percent change of –1.09 (95% CRI = [–2.11, –0.023]) over the entire time series 
and –3.51 (95% CRI = [–5.08, –1.88]) since 2013, when the population was listed 
as endangered. Given the magnitude of the decline, identifying which of the 
many factors affecting this population is most responsible is key in order to 
guide potential man­agement responses.

HŌ‘ULU‘ULU MANA‘O: Koho mākou i ka nui o nā pū‘uo ‘anehalapohe o nā koholā 
‘āhuka iwi poʼo like (Pseudorca crassidens) a puni ka paeʼāina ‘o Hawai‘i (MHI) 
mai nā makahiki 1999-2022 ma o kekahi ‘ōnaehana e kālailai ana i ka loa‘a ‘ana 
o ka holoholona i kekahi kālailai loa‘a-loaʼa hou. Mai nā makahiki 1999-2022 
mai, he mau ki‘ina ‘ohi hāpana i nānā ‘ia a ‘o ka loa‘a, ‘ike kūmakahiki ‘ia he 
265 koholā iwi po‘o like a he 53 mea i poelele lepili ‘ia. Kālailai pākahi ‘ia 
ke anapū‘uo a me ka ‘ikepili henua ma o nā koho pa‘apū kenele, a ‘o ka nui o ke 
kaulapa ma waena o ia mau ‘elua, ua ho‘ohana ‘ia i kumu ho‘ohālike i ka papaha 
‘ikena ma ka pū‘uo ho‘ohālike Bayesian Jolly-Sever. ‘Oi aku ka maika‘i o kēia 
hana ma mua o nā hana loa‘a-loa‘a hou ma‘amau no ka nānā ‘ana i nā ‘ano ao 
henua ‘ohi hāpana, a he ho‘oikaika i ke koho i ka ‘aui ‘ana o nā lawa kūpono. 
Ma ka ho‘ohana ʼana i ka ‘ikepili ho‘okūkohukohu ‘ia, he kumu ho‘ohālike 
maika‘i nō kēia i ka nui ‘ohi hāpana a me ka nui o nā hopena kaiaola, e la‘a 
ho‘i me ke kumuloli ‘ikena ha‘aha‘a, ka lō‘ihi o ka lepili ‘oko‘a, a me ke 
kumuloli o ka nui pū‘ulu launa. Ma ka launa ‘ana o ia kumu ho‘ohālike me ka 
‘ikepili o ko Hawai‘i pae‘āina koholā ‘āhuka iwi po‘o like, ‘ike mākou i ka 
li‘ili‘i mau o ka pū‘uo koholā ‘āhuka iwi po‘o like ‘ane‘ane puni ‘ia e ke kai, 
ma kahi o 139 (95% CRI = [114, 162]) ma ka makahiki 2022. Ma o ka wā kālailai, 
kohu mea lā, emi mai ka pū‘uo, he -1.09 pākēneka loli (95% CRI = [-2.11, 
-0.023]) ‘awelike o ka makahiki, a he -3.51 (95% CRI = [-5.08, -1.88]) mai ka 
makahiki 2013, i ka wā i helu ‘ia ai ka pū‘uo he mea ‘anehalapohe. No ka nui o 
ia ‘ano emi ‘ana, ko‘iko‘i nō ka ‘ike ‘ana i nā mea e pā nui ana i kēia pū‘uo i 
mea e alaka‘i aku ai i nā hāpane ho‘oponopono.

There is also a feature story on this publication on the NOAA Fisheries website 
at 
https://www.fisheries.noaa.gov/feature-story/endangered-population-false-killer-whales-main-hawaiian-islands-declining



--
Robin W. Baird, Ph.D.
Hawai‘i Program Director
Cascadia Research Collective<https://cascadiaresearch.org/>
218 ½ W. 4th Avenue
Olympia, WA 98501 USA
Donate to support our work<https://cascadiaresearch.org/donations/>
Follow us on Facebook<https://www.facebook.com/CascadiaResearch/>
Updates on our June O‘ahu field 
project<https://cascadiaresearch.org/hawaii-update/june2025/>

Recent publications on our 
work<https://cascadiaresearch.org/hawaii_publications/>:

Badger, J.J., R.W. Baird, D.S. Johnson, A.L. Bradford, S.D. Mahaffy, M.A. 
Kratofil, T. Cullins, J.J. Currie, S.H. Stack, and E.M. Oleson. 2025. 
Accounting for sampling bias reveals a decline in abundance of endangered false 
killer whales in the main Hawaiian Islands. Endangered Species Research 
https://doi.org/10.3354/esr01423

Baird, R.W. 2025. Out of sight, out of mind: false killer whale populations are 
at 
risk<https://cascadiaresearch.org/files/Baird_2025_Pseudorca-conservation_Whalewatcher.pdf>.
 Whalewatcher 45(1):76-82.

Corsi, E., R.W. Baird, A.E. Harnish, A.M. Gorgone, J.J. Currie, S.H. Stack, and 
J.J. Kiszka. 2025. Variation in social structure among multiple stocks of 
island-associated common bottlenose dolphins (Tursiops truncatus) in Hawaiian 
waters. Marine Mammal Science e70051. https://doi.org/10.1111/mms.70051

Henderson, E.E., M.A. Kratofil, R.W. Baird, C.R. Martin, A.E. Harnish, G.C. 
Alongi, S.W. Martin, and B.L. Southall. 2025. Exposure and response of 
satellite-tagged Blainville’s beaked whales to mid-frequency active sonar off 
Kaua‘i, Hawai‘i. Movement Ecology 13:29. 
https://doi.org/10.1186/s40462-025-00550-9

Mahaffy, S.D., U. Gorter, and R.W. Baird. 2025. Neonatal facial pigmentation of 
pygmy killer whales (Feresa attenuata): a description from repeated 
observations of individuals in the wild. Aquatic Mammals 51:138-143. 
https://doi.org/10.1578/AM51.2.2025.138

Walker-Milne, N.L., Y.P. Papastamatiou, S.D. Mahaffy, and R.W. Baird. 2025. 
Dynamics of foraging interactions between cookiecutter sharks (Isistius spp). 
and short-finned pilot whales (Globicephala macrorhynchus) in Hawai‘i. Marine 
Biology 172:63. https://doi.org/10.1007/s00227-025-04633-4

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