Yes!  I can't seem to find a copy of the article.  But going on your 
description and the figures, it looks like an excellent example of treating 
hierarchy as something to measure rather than impute. (The silverchair.com link 
didn't work, unfortunately.)

Until I can find a copy, some of what you say is provocative. It seems to me that talking 
directly about the graph (or network, an alternative Potochnik mentions) is the more 
literal concept, where level and hierarchy are the more metaphorical ones. Even the 
concept of accretion (temporal layering) is, to me, more meaningful than level or 
hierarchy.  So, the question remains *what* advantage do we gain from "zooming 
out" and thinking in terms of hierarchy and levels that we didn't already have in 
terms of [a]cyclic, temporal or structural, graphs?  Is the advantage largely rhetorical 
and communicative, accounting for the variations in the way the audience and participants 
think? Or are there, eg experimental design, questions and measures we can take that are 
made more precise and testable in terms of level and hierarchy versus graphs?

On 4/24/19 4:51 PM, Eric Smith wrote:
Here is a nice example, of that onus accepted and handled clearly.

https://www.nature.com/articles/nature07749
Topic is the accretionary dependency structure in the large subunit of the 
ribosome.

In particular, see Fig. 2, which my image-page on chrome is showing me at this 
URL (don’t know if these URLs produce equivalent output for different users):
https://www.google.com/search?q=bokov+and+steinberg+ribosome&source=lnms&tbm=isch&sa=X&ved=0ahUKEwjXjfm58unhAhXKzLwKHXG5B60Q_AUIDigB&biw=1371&bih=745#imgrc=uExkhZIl02WciM:

The primitive data is a set of links between locations in folded RNA, which can 
be assigned a directionality that is very likely a dynamically meaningful one.  
The result is a graph with directed links.  It is an empirical question whether 
the graph is cyclic or acyclic, with the answer being the latter.  The 
primitive data structure is only the acyclic graph.  However, a second question 
is whether the nodes in the graph admit a partial order, and if so, which sets 
of nodes constitute each distinct level within that order.  That question too 
has an answer in terms of the maximal extent to which the equivalence class 
defining a level can be extended, without violating the dependency structure in 
the underlying DAG.  Nodes in a level need not have been historically 
contemporaneous, but they reflect assembly conditions, as nodes at higher 
levels “plug into” nodes at lower levels, and thus require them to be in place. 
 This seems extremely likely to reflect an actual historical accretionary 
sequence, in which equivalence of nodes within a level quantifies the ambiguity 
of how they may have related in time.

Lots more has been done to extend this data to a detailed module decomposition, 
with or without the level post-processing.  Through all of it, the level 
decomposition continues to be salient, as levels by the analysis of the DAG 
also correspond roughly to horizons for generations of peptide structure.  See

https://watermark.silverchair.com/msx086.pdf?token=AQECAHi208BE49Ooan9kkhW_Ercy7Dm3ZL_9Cf3qfKAc485ysgAAAjwwggI4BgkqhkiG9w0BBwagggIpMIICJQIBADCCAh4GCSqGSIb3DQEHATAeBglghkgBZQMEAS4wEQQMGr_TvxBlD6v5A3yIAgEQgIIB77pRGYntr9gP-GNtZajC6JIiEDLCsmZFdcSgAVoYO43dh_vul542Uzn2GyejvMgnqthKt7u3ZnQoenITwMrwvneJMWZ9n6-UlYuottaxIkpxp6lWIfiTIla83YKJqigjdIbWtQx_W2y2J2pJgAKOBdbvvTctto3COkdwh4C6VH5AARmbw0bRfaMH_gRW8IKRNw8m4Gw--SbRMDlkHqaXRY8WJlbkrN8uB-ygTiu4TL12LHhNiWlxCLH0LP3pLKPBMmBG0tKM5sMIuO2CDVltBItUIT6i91Z0q2x-l6u5yBWqPFlDfpYNok--att5kqPbtzT1H7IzZev-AsWYpq_ek2RdyHxrthXdn2rTzvhMjmUlb1JHoeJX6holXrs8j1PKzwg_pW-3wtR6cYZg3VBLM6V_cTnMlyNIMABBkyix8D9pBvq6Hj7zLWABE8Oq0nuVUH5vd0U8RVbqpF5SS1OKd2Y13BN_bq-4P7B3RKKYmoecn2SVqoYPHZBV7csmkq9duwoydMQFbcGsk8BYopz6zEti3BuZJxXa2J6YT1i1pXQNMvSTHXRKdsIntCJkSZsPRwS-q6GiM5r7BtTU9hOLZLq__67NMjBDpWUcOG7pglEYuqENH7xy4abOEoE5TusJg9aU6PE9Tj9ayBkHnIONBg
and Fig.5 within it:
https://www.google.com/search?q=bokov+and+steinberg+ribosome&source=lnms&tbm=isch&sa=X&ved=0ahUKEwjXjfm58unhAhXKzLwKHXG5B60Q_AUIDigB&biw=1371&bih=745#imgrc=HRSn_FYi9cUYDM:

It’s great when people take on small enough questions that they have time to 
speak in full sentences.
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