I'm using Xubuntu 14.04. I solves the problem by setting 
/Net/EnableInputFeedbackSounds and /Net/EnableInputFeedbackSounds to true. By 
default their values are false. You can do this in setting manager or in 
terminal.
xfconf-query -c xsettings -p /Net/EnableEventSounds -s true
xfconf-query -c xsettings -p /Net/EnableInputFeedbackSounds -s true
I'm not sure if this applies to other Ubuntu flavours.

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https://bugs.launchpad.net/bugs/1492208

Title:
  No notification sound in thunderbird and lightning

Status in thunderbird package in Ubuntu:
  New

Bug description:
  14.04.3 TB 38.0.2
  Neither for new mails nor events

  I checked Sound effects aren't muted in control center, and main sound
  too isn't too low to be audible.

To manage notifications about this bug go to:
https://bugs.launchpad.net/ubuntu/+source/thunderbird/+bug/1492208/+subscriptions

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