This, along with https://bugs.launchpad.net/gnome-shell/+bug/1876628 occurred because I was using xserver-xorg-input-synaptics, which I gather is no longer supported. Removing this solved the problem.
-- You received this bug notification because you are a member of Ubuntu Desktop Bugs, which is subscribed to gnome-shell in Ubuntu. https://bugs.launchpad.net/bugs/1876630 Title: Application menu doesn't honour natural scroll on touchpad To manage notifications about this bug go to: https://bugs.launchpad.net/ubuntu/+source/gnome-shell/+bug/1876630/+subscriptions -- desktop-bugs mailing list desktop-bugs@lists.ubuntu.com https://lists.ubuntu.com/mailman/listinfo/desktop-bugs