upstream comments say:

The text entry for mail and web browsers is gone in GNOME 3. If you want to
setup another mail client than the ones listed, you'll need to create and
install a .desktop file to launch the application, which handles the
x-scheme-handler/mailto mime-type.

See how it was done for Evolution for example:
http://git.gnome.org/browse/evolution/commit/?id=61999b493fc4266300eab2907909867e73e4cc1e

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https://bugs.launchpad.net/bugs/416695

Title:
  No Browse button for Preferred Applications Mail Reader command

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